While Mr. Gold attempts to cross the Storybrooke border, Hook shoots Belle, causing her to fall over the border and lose her Enchanted Forest memories. As Mr. Gold is about to kill Hook by use of magic, a car runs into Storybrooke and hits Hook before swerving into a tree. Sheriff Emma Swan comes by with David and Mary Margaret to check out the site of the accident, and the trio is astonished to discover a stranger has made his way into Storybrooke from the outside. ("The Outsider")
Dr. Whale performs surgery on the stranger, whose phone Emma hacks into and learns his name is Greg Mendell. After the surgery is a success and Greg is resting up, Sheriff Emma questions him on what he saw before the car crashed. Greg casually notes he was looking down at his phone when the car got out of control, so he did not actually see anything. However, when Emma leaves his room, he reaches for his phone and makes a call to a person, saying, "You won't believe what I saw". ("In the Name of the Brother")
While still resting up in the hospital from his injuries, he visits Belle in her room. Greg explains he overheard what she said about having seen a man with a fireball in his hand. Belle thinks he is here to chastise her for her wild imagination, but instead believes her because he saw the same thing as well. ("Tiny")
Greg recovers well enough from his injuries to be up and walking around. While communicating via phone with the same unknown person, Greg says he wants to stay in town a little longer. When asked why, he proceeds to send a video of the town mayor, Regina, using a spell to summon the objects out of an unconscious Belle's purse. ("Manhattan")
Greg makes a trip to the diner, and asks that his order be packed to go, which the waitress Ruby wraps in a bag for him. He explains to Emma he is going hiking in the forest as per Dr. Whale's doctorly advice, and has an interest in nature photography. Emma thinks he would have gone back to Pennsylvania by now, but the town is beginning to grow on Greg.
In the woods, Greg bumps into Henry, and states he is taking a nature walk and snapping some photos. Henry points out the hiking trail is over that way, and Greg acts like he just found that out. Soon after, he makes a concerned phone call to Regina notifying her that Henry is by himself and seems upset. When Mary Margaret goes to Regina's doorstep to talk, he secretly records them from afar. He captures on film Regina ripping Mary Margaret's heart out, and then proceeding to put it back into her chest minutes later. Heading back into his car, he repeats the same words he said from childhood in vowing to find his father, and proceeds to turns on the ignition key, which has the woven leather key chain his father once gave him. ("Welcome to Storybrooke")
Shortly afterwards, he spends time at the diner eating apple pie. At his table booth, Regina formally introduces herself to Greg. Though she gets the feeling they have met before, he quickly dismisses her suspicions. No sooner had he returned to his hotel room, Greg meets Regina again, who now knows he is Owen. Greg demands to know what happened to his father, to which Regina explains Kurt was allowed to leave soon afterwards. Greg does not believe her, threatening to not leave until he finds his father. However, he is given a counter threat—contrary to what he believes, people can vanish into thin air. He later meets with "Her", who turns out to be Tamara, Neal Cassidy's fiancée; who is actually his secret lover. ("Selfless, Brave and True")
Greg shows Tamara on a map all the places he saw magic in Storybrooke. She asks him if he had any luck finding his dad. Greg says he hasn't found him but knows he is there. Then, before Tamara leaves, he asks her if the "package" is ready. She replies that she will be bringing it into town tonight.
Later in the evening, Greg meets Tamara by the town border where she drives a truck in with the package inside. He asks how the package would help and Tamara says that if there's anyone you could count on to do dirty work, it's a pirate. Then, she lifts the trunk revealing a captive and bound up Hook. ("Lacey")
With unholy intentions of stealing the magic from Storybrooke and finding his father, Greg and Tamara hold Hook captive in the clock tower. Hook questions why he was kidnapped and they explain if he helps them, they will assist him in killing the Dark One. Hook assures them he had already succeeded, but both prove him wrong and show him, through a telescope, a well and alive Mr. Gold. Hook is shocked and doubts whether the outsiders could help him.
Hook visits Regina at her office and warns her about the outsiders and their plans. Regina, who has discovered a method of escaping Storybrooke, clarifies to Hook he will get what he wants—the death of Mr. Gold—and be able to escape with her and Henry from the total destruction of Storybrooke, in return for his cooperation.
After Regina's successful attempt of acquiring the fail-safe trigger, she finds Hook alive in the library. Hook admits he has betrayed her to help Greg and Tamara in achieving their goal. Regina tries to use her magic against them, but fails. Greg explains her mother's glove—which Hook gave to her in the bottom of the library—was cuffed with the most powerful metal that man has ever known, which is why Regina was rendered powerless. When Tamara searches for Regina's identity through her list of Storybrooke residents and their true identities, and Greg states she is the Queen. Then, Greg commands Tamara to bag Regina. ("The Evil Queen")
Greg and Tamara kidnap and take Regina to the town's cannery storeroom where they strap her to a table and hook wires to connect her to the electrocution machine. Greg asks for Hook's help with the process, but Hook is not interested and curtly says he will be back when they want to help him kill Rumplestiltskin. Left alone with Regina, he takes great pleasure in turning the machine on and sending high powered electricity waves to shock Regina's body to get her to reveal his father's whereabouts.
Unknown to Greg and Tamara, Mary Margaret seeks out a way to gain a bodily connection with the missing Regina, and begins experiencing the same physical pain she is feeling from being electrocuted over and over. David reports to Emma and Neal, who are also on the lookout for Regina, of what Mary Margaret can remember from the momentarily connection. Upon mention of a strange smell of sardines in the area Regina is trapped in, Emma realizes they have to go to the cannery.
Tamara alerts Greg that Emma is onto them, and is closing in on the room, but he refuses to stop the procedure with Regina until he gets an answer out of her. He gives Regina one last chance before he turns the machine's power to the max. Finally, she admits to killing Kurt right after Greg left Storybrooke, and buried him at their former campsite. Then, she goads him into killing her himself. In his rage, Greg attempts to do just that, and shocks Regina until she passes out from the physical toll. He is forced to make his getaway after David bursts in and fires his gun.
Greg rushes off to the woods, and begins to shovel away the soil at the campsite location. When he hits something solid, desperation overtakes him and he uses his hands to claw away the dirt. His hands touch clothes he recognizes as his father's, and is saddened to see a skull. As he is reburying his father's remains, Tamara regroups with him. She gives her condolences about his father, and then shows him the diamond shaped item—a trigger—Regina had. Tamara sends it to be analyzed by the people they work for, and she says they need to take the next step by using it to destroy magic in Storybrooke. ("Second Star to the Right")
To use as a distraction, Tamara and Greg set off the trigger to buy themselves time in redirecting the whole town's attention onto the incoming catalyst. Desperate to save his own life, Hook aligns himself away from Tamara and Greg and gives away their location in helping David to steal back the remaining beans. David and Hook follow Greg into the cannery building. Inside, David threatens to pull the trigger unless the beans are handed over. As Greg pulls out a small clear bottle containing them, Tamara shoots as an ambush attack, though David manages to dodge while getting grazed by the bullet. Hook tackles Greg to the ground; causing the bottle to break on the floor. While David chases after Tamara, Hook wrestles for the beans and eventually grabs one of them while Greg makes off with the other.
During Tamara's getaway from David, she trips and her gun slides across the ground. Tamara is left vulnerable, but only for a short time, when Greg pushes her assailant down from behind. She motions to grab David's fallen gun, but her partner urges Tamara to go as they already got what they needed.
In the mines, while Emma figures out the trigger can be stopped if she combines her powers with Regina's, an impact blow in deactivating it causes everyone present to be thrown back onto the ground. Sometime during this time frame, Greg and Tamara make off with kidnapping Henry.
Outside, he and Tamara drag Henry towards the harbor. She ensures Henry they will not hurt him, to which he replies they will hurt everyone he loves, and just tried to destroy his hometown. In turn, Tamara counters back it was never their true intention. Greg reveals they wanted to put an end to magic, but found something much more valuable--him. Though Henry's family rush to save to stop them, they are not fast enough. The duo use their last magic bean to open a portal to Neverland; prompting the twosome to force Henry to jump in with them. ("And Straight On 'Til Morning")
Landing on the shores of Neverland's island, Henry attempts to make a hasty escape, but is swiftly caught by a water soaked Greg. Gleefully, Tamara remarks their mission has been accomplished, though Henry threatens that both of his mothers will be coming to save him. Greg dusts off his hopes by pointing out this new world is not anything like Storybrooke, and when Henry insists his family has been to the Enchanted Forest before, Tamara makes it known they are actually in Neverland. While she tries to use a communicator to let their employers, the Home Office, know they are present, Henry badgers Greg about who they work for and how they will get back home after magic is destroyed. To both questions, Greg remains tight-lipped, and stresses they don't need to know much about the Home Office, and believing in their cause alone is enough. Tamara interrupts to bring attention to the communicator's ineffectiveness, to which Greg opens the battery case to discover it's filled with sand. Shocked, she murmurs in disbelief of the current situation. Henry interjects it's good they don't ask any questions; causing Tamara to feel doubtful of the Home Office, but Greg presses on into the jungle where he starts a campfire as a signal. With what occurred earlier still fresh in Tamara's mind, she wonders if the empty communicator wasn't an accident, but Greg just thinks Henry's previous words are simply getting to her head.
Hearing nearby rustling, the brush parts to reveal a group of boys led by Felix. Greg demands to know who they are, and Felix states they are the Home Office. Unnerved, Tamara can't believe they've been working for are a bunch of teenagers, but Henry corrects her to say they are Lost Boys. He asks them why they want to destroy magic, but Felix acknowledges that was never their intention. Stunned, Greg says that was their mission, though Felix deflects the statement by countering that's what he and Tamara were told, and asks for Henry to be handed over. Refusing to comply, Tamara wants answers about the actual plan for magic and getting home, but is told they won't be going anywhere. Upset at being duped, Greg resists giving Henry over to them. With an ease, Felix, the lead Lost Boy, calls upon the Shadow, who rips out Greg's shadow; killing him. ("The Heart of the Truest Believer")